SearcharxivSearch

arXiv · 2110.01192

Influence of Allee Effect on Extreme Events in Coupled Three Species Systems

Abstract

We consider the dynamics of two coupled three-species population patches, incorporating the Allee Effect, focussing on the onset of extreme events in the coupled system. First we show that the interplay between coupling and the Allee effect may change the nature of the dynamics, with regular periodic dynamics becoming chaotic in a range of Allee parameters and coupling strengths. Further, the growth in the vegetation population displays an explosive blow-up beyond a critical value of coupling strength and Allee parameter. Most interestingly, we observe that beyond a threshold of coupling strength and Allee parameter, the population densities of all three species exhibit non-zero probability of yielding extreme events. The emergence of extreme events in the predator populations in the patches is the most prevalent, and the probability of obtaining large deviations in the predator populations is not affected significantly by either the coupling strength or the Allee effect. In the absence of the Allee effect the prey population in the coupled system exhibits no extreme events for low coupling strengths, but yields a sharp increase in extreme events after a critical strength of coupling. The vegetation population in the patches display a small finite probability of extreme events for strong enough coupling, only in the presence of Allee effect. Lastly we consider the influence of additive noise on the continued prevalence of extreme events. Very significantly, we find that noise suppresses the unbounded vegetation growth that was induced by a combination of Allee effect and coupling. Further, we demonstrate that noise mitigates extreme events in all three populations, and beyond a noise level we do not observe any extreme events in the system at all. This finding has important bearing on the potential observability of extreme events in natural and laboratory systems.

Explore related subjects

Keep this discovery

BibTeXRIS

Deeptajyoti Sen, Sudeshna Sinha. 2021-10-04. Influence of Allee Effect on Extreme Events in Coupled Three Species Systems. https://arxiv.org/abs/2110.01192

Cite the original work for its findings. Save a collection to share your selection of sources.

KEEP EXPLORING

Related papers

Competition drives excessive recruitment in collective search

Groups that search collectively often exploit what they find by recruiting: one member directs others to a site it has found. Recruitment raises the number of members foraging at a known site, but the return per forager may fall as that number grows, so there is an intermediate optimal recruitment rate. In addition, a site may be used by more than one group. Here we analyze a model of two groups that forage from a single site whose return declines with the total number of foragers present. The two groups interact only through this shared return. The long-run outcome is either coexistence at the foraging site or monopoly by one group, and we analyze the boundary between these two outcomes. A group's best response to its rival is not monotone: it increases its own recruitment rate with the rival's recruitment rate in an attempt to preserve a monopoly, and then its recruitment rate drops discontinuously when it is no longer optimal to preserve a monopoly. We analyze how model parameters govern this shift: a group relinquishes monopoly when the site saturates at few foragers and when the rival group is small. When the two groups have comparable size there are multiple Nash equilibria, so either group may end up with the larger share. And when two equally matched groups compete, both recruit above the rate that maximizes their common return, so that each individual ends with less than it would in a single undivided group of the same total size.

q-bio.PE

Selection Rules for Species Coexistence in a Hierarchical May-Leonard Model

One of the central challenges in evolutionary dynamics is understanding why some species combinations persist while others disappear. Although cyclic-interaction models have provided fundamental insights into biodiversity maintenance, much less is known about how hierarchical competitive interactions shape long-term community organization. Here, we investigate a hierarchical extension of the May-Leonard model, in which species interact through a directed predation chain while undergoing reproduction and mortality. Combining mean-field analysis with Monte Carlo simulations, we show that the fully coexisting state is generically unstable, causing the dynamics to evolve toward lower-dimensional coexistence states. The simulations further reveal stochastic extinctions dominating small populations with the dynamics progressively approaching the mean-field predictions as the system size increases. Rather than permitting arbitrary species combinations, the hierarchical-interaction structure dynamically constrains coexistence by selecting only specific subsets of species for long-term persistence. We show that these admissible coexistence states have a natural graph-theoretic interpretation as independent sets in the hierarchical interaction network, thereby providing general constraints on coexistence in hierarchical communities. Together, these results establish a theoretical framework linking hierarchical interactions, dynamical selection, graph topology, and biodiversity organization, extending the classical May-Leonard model beyond cyclic competition.

q-bio.PE

Persistence of n-Species Lotka-Volterra Models with Periodic Pulses

Periodic impulsive interventions arise naturally in the management of biological populations, including chemotherapy, pesticide application, and infectious-disease treatment. We develop general conditions for permanence in n-species population models subject to periodic multiplicative pulse disturbances. Our main result provides a sufficient condition for permanence in terms of weighted long-term growth rates on a Morse decomposition of the extinction set, explicitly separating the contributions of continuous population dynamics from those of the periodic pulse. To establish this result, we transform the impulsive system into an associated autonomous continuous-time dynamical system and use this correspondence to extend classical permanence theory to periodically pulsed models. We further show that the same conditions imply robust permanence under sufficiently small perturbations to the continuous dynamics, pulse period, and pulse effects. We illustrate the framework with two Lotka-Volterra models motivated by biological control: competition between chemotherapy-sensitive and chemotherapy-resistant cancer cells, and integrated control of an agricultural pest using pesticides and parasitoids. These examples demonstrate how intervention frequency and intensity interact with underlying ecological interactions to determine whether populations coexist or are excluded. Our results provide a general framework for analyzing persistence in ecological systems subject to repeated discrete disturbances.

q-bio.PE