arXiv subjects

Alexandru I. Tomescu

Publications and source records attributed to Alexandru I. Tomescu.

At least 19 recordsLinked to original sources

arXiv2026

Fast and Flexible Flow Decompositions in General Graphs via Dominators

Multi-assembly methods rely at their core on a flow decomposition problem, namely, decomposing a weighted graph into weighted paths or walks. However, most results over the past decade have focused on decompositions over directed acyclic graphs (DAGs). This limitation has lead to either purely heuristic methods, or in applications transforming a graph with cycles into a DAG via preprocessing heuristics. In this paper we show that flow decomposition problems can be solved in practice also on general graphs with cycles, via a framework that yields fast and flexible Mixed Integer Linear Programming (MILP) formulations. Our key technique relies on the graph-theoretic notion of dominator tree, which we use to find all safe sequences of edges, that are guaranteed to appear in some walk of any flow decomposition solution. We generalize previous results from DAGs to cyclic graphs, by showing that maximal safe sequences correspond to extensions of common leaves of two dominator trees, and that we can find all of them in time linear in their size. Using these, we can accelerate MILPs for any flow decomposition into walks in general graphs, by setting to (at least) 1 suitable variables encoding solution walks, and by setting to 0 other walks variables non-reachable to and from safe sequences. This reduces model size and eliminates costly linearizations of MILP variable products. We experiment with three decomposition models (Minimum Flow Decomposition, Least Absolute Errors and Minimum Path Error), on four bacterial datasets. Our pre-processing enables up to thousand-fold speedups and solves even under 30 seconds many instances otherwise timing out. We thus hope that our dominator-based MILP simplification framework, and the accompanying software library can become building blocks in multi-assembly applications.

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arXiv2026

Maximum Coverage $k$-Antichains and Chains: A Greedy Approach

Given an acyclic digraph $G = (V,E)$ and a positive integer $k$, the problem of Maximum Coverage $k$-Antichains (resp. Chains) denoted as MA-$k$ (resp. MC-$k$) asks to find $k$ sets of pairwise unreachable vertices, known as antichains (resp. $k$ subsequences of paths, known as chains), maximizing the number $α_k$ (resp. $β_k$) of vertices covered by these antichains (resp. chains). While MC-$k$ was solved in almost optimal $|E|^{1+o(1)}$ time~[Kogan and Parter, ICALP'22], the fastest algorithms for MA-$k$ are a $(k|E|)^{1+o(1)}$-time solution and a $|E|^{1+o(1)}$-time $1/2$ approximation~[Kogan and Parter, ESA'24]. We obtain the following for MA-$k$: - An algorithm running in $|E|^{1+o(1)}$ time, and an algorithm running in parameterized near-linear $\tilde{O}(α_k |E|)$ time. Our algorithms are simple solutions exploiting a paths-based proof of the Greene-Kleitman theorems leveraged by the greedy algorithm for set cover as well as recent advances in fast algorithms for flows and shortest paths. - An approximation algorithm running in parameterized linear time $O(α_1^2|V| + (α_1+k)|E|)$ with approximation ratio of $(1-1/e) > 0.63 > 1/2$, beating the state-of-the-art $1/2$ approximation. Our solution uses greedy for antichains and a simple strategy to amortize the cost of computing consecutive maximum antichains. We complement these results with two examples (one for chains and one for antichains) showing that, for every $k \ge 2$, greedy misses the tight $1/e$ portion of the optimal coverage for chains, and a $1/4$ portion for antichains. We also show that greedy is a $Ω(\log{|V|})$ factor away from minimality when required to cover all vertices: previously unknown for sets of chains or antichains.

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arXiv2026

The Power of Graph Doubling: Computing Ultrabubbles in a Bidirected Graph by Reducing to Weak Superbubbles

Bidirected graphs are a common generalisation of directed graphs where arcs can also be incoming to both their incident nodes, or outgoing from both their incident nodes. Such arcs allow a walk to change direction. Some algorithms can easily be adapted from directed graphs to bidirected graphs, such as shortest path algorithms. These adaptions are already used in practice, and implicitly use the graph doubling technique to apply an algorithm for directed graphs to bidirected graphs. In other cases, the applicability of graph doubling is not that obvious. For example, superbubbles and their generalisation to bidirected graphs ultrabubbles. Ultrabubbles are a common structure in bidirected biological graphs which carries biological meaning, but also functions as a nested clustering method, since an ultrabubble is separated by only two nodes from the rest of the graph. There is an existing method that enumerates a structure similar to ultrabubbles by enumerating (weak) superbubbles in the doubled graph. However, the literature does not make any direct connection between superbubbles and ultrabubbles except that a superbubble is an ultrabubble in a directed graph. Only a partial result connecting superbubbles and ultrabubbles exists by Harviainen et al. (2026). Graph doubling on the other hand maintains connectivity, and allows to draw a direct connection between ultrabubbles and weak superbubbles. This results in the first linear-time reduction-based algorithm for computing ultrabubbles on any bidirected graph. Together with the fact that graph doubling is already used implicitly in simple cases, our result motivates that graph doubling is a powerful yet simple technique to apply algorithms for directed graphs to bidirected graphs.

cs.DS↗

arXiv2026

Identifying bubble-like subgraphs in linear-time via a unified SPQR-tree framework

A fundamental algorithmic problem in computational biology is to find all subgraphs of a given type (superbubbles, snarls, and ultrabubbles) in a directed or bidirected input graph. These correspond to regions of genetic variation and are useful in analyzing collections of genomes. We present the first linear-time algorithms for identifying all snarls and all ultrabubbles, resolving problems open since 2018. The algorithm for snarls relies on a new linear-size representation of all snarls with respect to the number of vertices in the graph. We employ the well-known SPQR-tree decomposition, which encodes all 2-separators of a biconnected graph. After several dynamic-programming-style traversals of this tree, we maintain key properties (such as acyclicity) that allow us to decide whether a given 2-separator defines a subgraph to be reported. A crucial ingredient for linear-time complexity is that acyclicity of linearly many subgraphs can be tested simultaneously via the problem of computing all arcs in a directed graph whose removal renders it acyclic (so-called feedback arcs). As such, we prove a fundamental result for bidirected graphs, that may be of independent interest: all feedback arcs can be computed in linear time for tipless bidirected graphs, while in general this is at least as hard as matrix multiplication, assuming the k-Clique Conjecture. Our results form a unified framework that also yields a completely different linear-time algorithm for finding all superbubbles. Although some of the results are technically involved, the underlying ideas are conceptually simple, and may extend to other bubble-like subgraphs. More broadly, our work contributes to the theoretical foundations of computational biology and advances a growing line of research that uses SPQR-tree decompositions as a general tool for designing efficient algorithms, beyond their traditional role in graph drawing.

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arXiv2026

EMERALD-UI: An interactive web application to unveil novel protein biology hidden in the suboptimal-alignment space

Life over the past four billion years has been shaped by proteins and their capacity to assemble into three dimensional conformations. Protein sequence alignments have been the enabling technology for exploring the evolution and functional adaptation of proteins across the tree of life. Recent advancements in scaling the prediction of three dimensional protein structures from primary sequence alone, revealed that different modes of conservation and function operate on the sequence and structure level. This difference in protein conservation patterns and their underlying functional change that could emerge in suboptimal alignment configurations is often ignored in optimal protein alignment approaches. We introduce EMERALD-UI, an open-source interactive web application which is designed to reveal unexplored biology by visualising stable structural conformations or protein regions hidden in the suboptimal alignment space. Availability: EMERALD-UI is available at https://algbio.github.io/emerald-ui/. Contact: hdrost001@dundee.ac.uk or alexandru.tomescu@helsinki.fi.

q-bio.QM↗

arXiv2025

Width Parameters for Minimum Flow Decomposition

Minimum flow decomposition (MFD) is the strongly NP-hard problem of finding a smallest set of integer weighted $s$-$t$ paths in an $s$-$t$ DAG $G$ whose weighted sum is equal to a given flow $f$ on $G$. Despite its many practical applications, we lack an understanding of graph structures that make MFD easy or hard. Recent progress is due to Cáceres et al. [ACM TALG 2024], who showed that the DAG width, the minimum number of paths to cover all edges, plays an essential role in the approximation of the problem. Our first set of results regard the computational complexity of MFD parameterised by the width. This question was previously open, because MFD on width-1 DAGs (paths) is trivially solvable, and the existing NP-hardness proofs use DAGs of unbounded width. We show that MFD on width-2 DAGs is already NP-hard and that MFD on width-3 DAGs is strongly NP-hard. Our main contribution complements these hardness bounds, as we show that weak NP-hardness is the best we can hope for on width-2 DAGs. In fact, we prove the more general statement that MFD with unary coded input can be solved in quasi-polynomial time on DAGs of constant parallel-width, which includes width-2 DAGs. The parallel-width of a DAG $G$ (par-width$(G)$) was defined by Deligkas and Meir [MFCS 2017] as the size of the largest minimal $s$-$t$ cut-set. We obtain these results by, a) interpreting flow decompositions as a sequence of certain digraph minor operations defined by Deligkas and Meir [MFCS 2017], and b) defining a new notion of width of a flow network, flow-width of $(G,f)$, defined as the minimum number of paths covering all edges of $G$, where every edge $e$ can be covered by at most $f(e)$ paths. Using (a) and (b), we show as an intermediate result, an improved upper bound $(\lfloor\log \Vert f\Vert\rfloor+1) \cdot \text{par-width}(G)$ for MFD, where $\Vert f\Vert$ is the largest flow weight of all edges.

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arXiv2025

Identifying all snarls and superbubbles in linear-time, via a unified SPQR-tree framework

Snarls and superbubbles are fundamental pangenome decompositions capturing variant sites. These bubble-like structures underpin key tasks in computational pangenomics, including structural-variant genotyping, distance indexing, haplotype sampling, and variant annotation. Snarls can be quadratically-many in the size of the graph, and since their introduction in 2018 with the vg toolkit, there has been no work on identifying all snarls in linear time. Moreover, while it is known how to find superbubbles in linear time, this result is a highly specialized solution only achieved after a long series of papers. We present the first algorithm identifying all snarls in linear time. This is based on a new representation of all snarls, of size linear in the input graph size, and which can be computed in linear time. Our algorithm is based on a unified framework that also provides a new linear-time algorithm for finding superbubbles. An observation behind our results is that all such structures are separated from the rest of the graph by two vertices (except for cases which are trivially computable), i.e. their endpoints are a 2-separator of the underlying undirected graph. Based on this, we employ the well-known SPQR tree decomposition, which encodes all 2-separators, to guide a traversal that finds the bubble-like structures efficiently. We implemented our algorithms in C++ (available at https://github.com/algbio/BubbleFinder) and evaluated them on various pangenomic datasets. Our algorithms outcompete or they are on the same level of existing methods. For snarls, we are up to two times faster than vg, while identifying all snarls. When computing superbubbles, we are up to 50 times faster than BubbleGun. Our SPQR tree framework provides a unifying perspective on bubble-like structures in pangenomics, together with a template for finding other bubble-like structures efficiently.

cs.DS↗

arXiv2025

Safe Sequences via Dominators in DAGs for Path-Covering Problems

A path-covering problem on a directed acyclic graph (DAG) requires finding a set of source-to-sink paths that cover all the nodes, all the arcs, or subsets thereof, and additionally they are optimal with respect to some function. In this paper we study safe sequences of nodes or arcs, namely sequences that appear in some path of every path cover of a DAG. We show that safe sequences admit a simple characterization via cutnodes. Moreover, we establish a connection between maximal safe sequences and leaf-to-root paths in the source- and sink-dominator trees of the DAG, which may be of independent interest in the extensive literature on dominators. With dominator trees, safe sequences admit an O(n)-size representation and a linear-time output-sensitive enumeration algorithm running in time O(m + o), where n and m are the number of nodes and arcs, respectively, and o is the total length of the maximal safe sequences. We then apply maximal safe sequences to simplify Integer Linear Programs (ILPs) for two path-covering problems, LeastSquares and MinPathError, which are at the core of RNA transcript assembly problems from bioinformatics. On various datasets, maximal safe sequences can be computed in under 0.1 seconds per graph, on average, and ILP solvers whose search space is reduced in this manner exhibit significant speed-ups. For example on graphs with a large width, average speed-ups are in the range 50-250x for MinPathError and in the range 80-350x for LeastSquares. Optimizing ILPs using safe sequences can thus become a fast building block of practical RNA transcript assembly tools, and more generally, of path-covering problems.

cs.DS↗

arXiv2023

Accelerating ILP solvers for Minimum Flow Decompositions through search space and dimensionality reductions

Given a flow network, the Minimum Flow Decomposition (MFD) problem is finding the smallest possible set of weighted paths whose superposition equals the flow. It is a classical, strongly NP-hard problem that is proven to be useful in RNA transcript assembly and applications outside of Bioinformatics. We improve an existing ILP (Integer Linear Programming) model by Dias et al. [RECOMB 2022] for DAGs by decreasing the solver's search space using solution safety and several other optimizations. This results in a significant speedup compared to the original ILP, of up to 55-90x on average on the hardest instances. Moreover, we show that our optimizations apply also to MFD problem variants, resulting in similar speedups, going up to 123x on the hardest instances. We also developed an ILP model of reduced dimensionality for an MFD variant in which the solution path weights are restricted to a given set. This model can find an optimal MFD solution for most instances, and overall, its accuracy significantly outperforms that of previous greedy algorithms while being up to an order of magnitude faster than our optimized ILP.

q-bio.GN↗

arXiv2023

Minimum Path Cover: The Power of Parameterization

Computing a minimum path cover (MPC) of a directed acyclic graph (DAG) is a fundamental problem with a myriad of applications, including reachability. Although it is known how to solve the problem by a simple reduction to minimum flow, recent theoretical advances exploit this idea to obtain algorithms parameterized by the number of paths of an MPC, known as the width. These results obtain fast [Mäkinen et al., TALG] and even linear time [Cáceres et al., SODA 2022] algorithms in the small-width regime. In this paper, we present the first publicly available high-performance implementation of state-of-the-art MPC algorithms, including the parameterized approaches. Our experiments on random DAGs show that parameterized algorithms are orders-of-magnitude faster on dense graphs. Additionally, we present new pre-processing heuristics based on transitive edge sparsification. We show that our heuristics improve MPC-solvers by orders-of-magnitude.

cs.DS↗

arXiv2023

Simplicity in Eulerian Circuits: Uniqueness and Safety

An Eulerian circuit in a directed graph is one of the most fundamental Graph Theory notions. Detecting if a graph $G$ has a unique Eulerian circuit can be done in polynomial time via the BEST theorem by de Bruijn, van Aardenne-Ehrenfest, Smith and Tutte, 1941-1951 (involving counting arborescences), or via a tailored characterization by Pevzner, 1989 (involving computing the intersection graph of simple cycles of $G$), both of which thus rely on overly complex notions for the simpler uniqueness problem. In this paper we give a new linear-time checkable characterization of directed graphs with a unique Eulerian circuit. This is based on a simple condition of when two edges must appear consecutively in all Eulerian circuits, in terms of cut nodes of the underlying undirected graph of $G$. As a by-product, we can also compute in linear-time all maximal $\textit{safe}$ walks appearing in all Eulerian circuits, for which Nagarajan and Pop proposed in 2009 a polynomial-time algorithm based on Pevzner characterization.

cs.DS↗

arXiv2023

Width Helps and Hinders Splitting Flows

Minimum flow decomposition (MFD) is the NP-hard problem of finding a smallest decomposition of a network flow/circulation $X$ on a directed graph $G$ into weighted source-to-sink paths whose superposition equals $X$. We show that, for acyclic graphs, considering the \emph{width} of the graph (the minimum number of paths needed to cover all of its edges) yields advances in our understanding of its approximability. For the version of the problem that uses only non-negative weights, we identify and characterise a new class of \emph{width-stable} graphs, for which a popular heuristic is a \gwsimple-approximation ($|X|$ being the total flow of $X$), and strengthen its worst-case approximation ratio from $Ω(\sqrt{m})$ to $Ω(m / \log m)$ for sparse graphs, where $m$ is the number of edges in the graph. We also study a new problem on graphs with cycles, Minimum Cost Circulation Decomposition (MCCD), and show that it generalises MFD through a simple reduction. For the version allowing also negative weights, we give a $(\lceil \log \Vert X \Vert \rceil +1)$-approximation ($\Vert X \Vert$ being the maximum absolute value of $X$ on any edge) using a power-of-two approach, combined with parity fixing arguments and a decomposition of unitary circulations ($\Vert X \Vert \leq 1$), using a generalised notion of width for this problem. Finally, we disprove a conjecture about the linear independence of minimum (non-negative) flow decompositions posed by Kloster et al. [ALENEX 2018], but show that its useful implication (polynomial-time assignments of weights to a given set of paths to decompose a flow) holds for the negative version.

cs.DS↗

arXiv2023

A Safety Framework for Flow Decomposition Problems via Integer Linear Programming

Many important problems in Bioinformatics (e.g., assembly or multi-assembly) admit multiple solutions, while the final objective is to report only one. A common approach to deal with this uncertainty is finding safe partial solutions (e.g., contigs) which are common to all solutions. Previous research on safety has focused on polynomially-time solvable problems, whereas many successful and natural models are NP-hard to solve, leaving a lack of "safety tools" for such problems. We propose the first method for computing all safe solutions for an NP-hard problem, minimum flow decomposition. We obtain our results by developing a "safety test" for paths based on a general Integer Linear Programming (ILP) formulation. Moreover, we provide implementations with practical optimizations aimed to reduce the total ILP time, the most efficient of these being based on a recursive group-testing procedure. Results: Experimental results on the transcriptome datasets of Shao and Kingsford (TCBB, 2017) show that all safe paths for minimum flow decompositions correctly recover up to 90% of the full RNA transcripts, which is at least 25% more than previously known safe paths, such as (Caceres et al. TCBB, 2021), (Zheng et al., RECOMB 2021), (Khan et al., RECOMB 2022, ESA 2022). Moreover, despite the NP-hardness of the problem, we can report all safe paths for 99.8% of the over 27,000 non-trivial graphs of this dataset in only 1.5 hours. Our results suggest that, on perfect data, there is less ambiguity than thought in the notoriously hard RNA assembly problem. Availability: https://github.com/algbio/mfd-safety

cs.DS↗

arXiv2023

Minimum Flow Decomposition in Graphs with Cycles using Integer Linear Programming

Minimum flow decomposition (MFD) -- the problem of finding a minimum set of weighted source-to-sink paths that perfectly decomposes a flow -- is a classical problem in Computer Science, and variants of it are powerful models in different fields such as Bioinformatics and Transportation. Even on acyclic graphs, the problem is NP-hard, and most practical solutions have been via heuristics or approximations. While there is an extensive body of research on acyclic graphs, currently, there is no \emph{exact} solution on graphs with cycles. In this paper, we present the first ILP formulation for three natural variants of the MFD problem in graphs with cycles, asking for a decomposition consisting only of weighted source-to-sink paths or cycles, trails, and walks, respectively. On three datasets of increasing levels of complexity from both Bioinformatics and Transportation, our approaches solve any instance in under 10 minutes. Our implementations are freely available at github.com/algbio/MFD-ILP.

cs.DS↗

arXiv2022

Minimum Path Cover in Parameterized Linear Time

A minimum path cover (MPC) of a directed acyclic graph (DAG) $G = (V,E)$ is a minimum-size set of paths that together cover all the vertices of the DAG. Computing an MPC is a basic polynomial problem, dating back to Dilworth's and Fulkerson's results in the 1950s. Since the size $k$ of an MPC (also known as the width) can be small in practical applications, research has also studied algorithms whose running time is parameterized on $k$. We obtain a new MPC parameterized algorithm for DAGs running in time $O(k^2|V| + |E|)$. Our algorithm is the first solving the problem in parameterized linear time. Additionally, we obtain an edge sparsification algorithm preserving the width of a DAG but reducing $|E|$ to less than $2|V|$. This algorithm runs in time $O(k^2|V|)$ and requires an MPC of a DAG as input, thus its total running time is the same as the running time of our MPC algorithm.

cs.DS↗

arXiv2022

Cut paths and their remainder structure, with applications

In a strongly connected graph $G = (V,E)$, a cut arc (also called strong bridge) is an arc $e \in E$ whose removal makes the graph no longer strongly connected. Equivalently, there exist $u,v \in V$, such that all $u$-$v$ walks contain $e$. Cut arcs are a fundamental graph-theoretic notion, with countless applications, especially in reachability problems. In this paper we initiate the study of cut paths, as a generalisation of cut arcs, which we naturally define as those paths $P$ for which there exist $u,v \in V$, such that all $u$-$v$ walks contain $P$ as subwalk. We first prove various properties of cut paths and define their remainder structures, which we use to present a simple $O(m)$-time verification algorithm for a cut path ($|V| = n$, $|E| = m$). Secondly, we apply cut paths and their remainder structures to improve several reachability problems from bioinformatics. A walk is called safe if it is a subwalk of every node-covering closed walk of a strongly connected graph. Multi-safety is defined analogously, by considering node-covering sets of closed walks instead. We show that cut paths provide simple $O(m)$-time algorithms verifying if a walk is safe or multi-safe. For multi-safety, we present the first linear time algorithm, while for safety, we present a simple algorithm where the state-of-the-art employed complex data structures. Finally we show that the simultaneous computation of remainder structures of all subwalks of a cut path can be performed in linear time. These properties yield an $O(mn)$ algorithm outputting all maximal multi-safe walks, improving over the state-of-the-art algorithm running in time $O(m^2+n^3)$. The results of this paper only scratch the surface in the study of cut paths, and we believe a rich structure of a graph can be revealed, considering the perspective of a path, instead of just an arc.

cs.DM↗

arXiv2022

Optimizing Safe Flow Decompositions in DAGs

Network flow is one of the most studied combinatorial optimization problems having innumerable applications. Any flow on a directed acyclic graph $G$ having $n$ vertices and $m$ edges can be decomposed into a set of $O(m)$ paths. In some applications, each solution (decomposition) corresponds to some particular data that generated the original flow. Given the possibility of multiple optimal solutions, no optimization criterion ensures the identification of the correct decomposition. Hence, recently flow decomposition was studied [RECOMB22] in the Safe and Complete framework, particularly for RNA Assembly. They presented a characterization of the safe paths, resulting in an $O(mn+out_R)$ time algorithm to compute all safe paths, where $out_R$ is the size of the raw output reporting each safe path explicitly. They also showed that $out_R$ can be $Ω(mn^2)$ in the worst case but $O(m)$ in the best case. Hence, they further presented an algorithm to report a concise representation of the output $out_C$ in $O(mn+out_C)$ time, where $out_C$ can be $Ω(mn)$ in the worst case but $O(m)$ in the best case. In this work, we study how different safe paths interact, resulting in optimal output-sensitive algorithms requiring $O(m+out_R)$ and $O(m+out_C)$ time for computing the existing representations of the safe paths. Further, we propose a new characterization of the safe paths resulting in the {\em optimal} representation of safe paths $out_O$, which can be $Ω(mn)$ in the worst case but requires optimal $O(1)$ space for every safe path reported, with a near-optimal computation algorithm. Overall we further develop the theory of safe and complete solutions for the flow decomposition problem, giving an optimal algorithm for the explicit representation, and a near-optimal algorithm for the optimal representation of the safe paths

cs.DS↗

arXiv2022

Algorithms and Complexity on Indexing Founder Graphs

We study the problem of matching a string in a labeled graph. Previous research has shown that unless the Orthogonal Vectors Hypothesis (OVH) is false, one cannot solve this problem in strongly sub-quadratic time, nor index the graph in polynomial time to answer queries efficiently (Equi et al. ICALP 2019, SOFSEM 2021). These conditional lower-bounds cover even deterministic graphs with binary alphabet, but there naturally exist also graph classes that are easy to index: E.g. Wheeler graphs (Gagie et al. Theor. Comp. Sci. 2017) cover graphs admitting a Burrows-Wheeler transform -based indexing scheme. However, it is NP-complete to recognize if a graph is a Wheeler graph (Gibney, Thankachan, ESA 2019). We propose an approach to alleviate the construction bottleneck of Wheeler graphs. Rather than starting from an arbitrary graph, we study graphs induced from multiple sequence alignments (MSAs). Elastic degenerate strings (Bernadini et al. SPIRE 2017, ICALP 2019) can be seen as such graphs, and we introduce here their generalization: elastic founder graphs. We first prove that even such induced graphs are hard to index under OVH. Then we introduce two subclasses, repeat-free and semi-repeat-free graphs, that are easy to index. We give a linear time algorithm to construct a repeat-free non-elastic founder graph from a gapless MSA, and (parameterized) near-linear time algorithms to construct semi-repeat-free (repeat-free, respectively) elastic founder graphs from general MSAs. Finally, we show that repeat-free elastic founder graphs admit a reduction to Wheeler graphs in polynomial time.

cs.DS↗