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Rana D. Parshad

Publications and source records attributed to Rana D. Parshad.

At least 19 recordsLinked to original sources

RevCRN: Reversible Analog Computation using Chemical Reaction Networks

The computability of real numbers and functions using Turing Machines has been a central area of theoretical computer science since the mid-20th century. In the late 20th century, it was shown that chemical reactions can serve as a basis for computation using the Chemical Reaction Network (CRN) model. Recent advances in computing real numbers using Deterministic Chemical Reaction Networks (DCRNs) have identified numerous classes of DCRN-computable real numbers. In parallel, the works of R. Landauer and C. H. Bennett, spanning the 1960s to the early 2000s, showed that reversible computing offers significant advantages over irreversible methods, particularly in energy efficiency, motivating extensive research on reversible computation. In this work, we investigate the computability of real numbers using Reversible Chemical Reaction Networks (RevCRNs). The paper has two primary contributions: (1) establishing relationships among CRN-computable real number classes including Lyapunov CRN ($\mathbb{R}_{LCRN}$), Real-Time CRN ($\mathbb{R}_{RTCRN}$), rational numbers ($\mathbb{Q}$), and RevCRNs ($\mathbb{R}_{RevCRN}$), with key results: (i) $\mathbb{Q}$ is a strict subset of $\mathbb{R}_{RevCRN}$; (ii) the set of positive algebraic numbers ($ALG$), $\mathbb{R}_{LCRN}$, and real numbers computable by 1-species RevCRN ($\mathbb{R}_{RevCRN}^{1s}$) are equal; (iii) $\mathbb{R}_{RTCRN}$ and $\mathbb{R}_{RevCRN}$ exhibit non-empty overlap; and (iv) the set of real numbers computable by detailed-balanced RevCRNs ($\mathbb{R}^{DetBal}_{RevCRN}$) is a subset of $ALG$; and (2) exploring the existence of a hierarchy within $\mathbb{R}_{RevCRN}$. Finally, we leave open the exact relationship between $\mathbb{R}_{RevCRN}$ and $\mathbb{R}_{RTCRN}$ while conjecturing a general hierarchy of RevCRN-computable reals.

cs.CC

Novel Dynamics in Models of Angiogenesis with p-Laplacian diffusion

Ischemic heart diseases represent the leading cause of mortality worldwide. Revascularization, the process to restore blood flow in blockages, shows promise. To this end, mathematical models for angiogenesis, the process by which new blood vessels form from existing ones, have been extremely well investigated. In the current work, we consider a classical two species model for angiogenesis, consisting of cell and VEGF populations. However, we assume the cells move according to p-Laplacian diffusion, which could be both ``fast" ($1 2$), in addition to normal diffusion ($p=2$). We first show that the system is well posed in a weak sense when $p>\frac{3}{2}$, for sufficiently small initial data. Next, we show that the p-Laplacian can lead to several novel dynamics not reported earlier; these include increased cellular proliferation via bi-modal and multi spike solutions, gain of regularity, prevention of finite time blow-up, cell depletion via finite time extinction, and Turing patterns. We discuss applications of these results for cardiac health via a digital twins framework.

math.AP

Additional Food Enhances the Bifurcation Structure of Predator Competition Models

Additional food sources and predator competition are both known to impact the dynamics of predator-prey models. The Bazykin model of predator competition, with Holling type II functional response, possesses a rich bifurcation structure consisting of a focus-type degenerate Bogdanov-Takens bifurcation of codimension 3, and a degenerate Hopf bifurcation of codimension at most 2. Additional food models on the other hand are able to drive pest populations lower, with vast applicability in biological control. Despite these models being studied rigorously in the literature, the global bifurcation structure, of their possible complex dynamics, in a unified model, is unknown. In this work, we study an additional food model with generalized predator competition and Holling type-II functional response. Depending on the parameter values, the system can have up to three interior equilibria. Further, we show that this system exhibits a cusp-type (or focus-type) Bogdanov-Takens bifurcation of codimension at least 4 (or 3), a global Hopf bifurcation of codimension 3, and a homoclinic bifurcation of codimension 3. This shows there could exist three limit cycles around the BT point. These results demonstrate that additional food in Bazykin type models, can enhance their bifurcation structure. We discuss the applicability of these results to integrated pest management programs for the soybean aphid, wherein long term field data in the North-Central United States, shows two distinct limit cycles in aphid populations and their predators. Our results suggest biological control with additional food, is an effective management tactic for invasive pests.

q-bio.PE

An Investigation of Additional Food Models with Generalised Functional Response

Additional food sources are often used to improve the effectiveness of predators in controlling pest populations. However, the non-symmetric structure of additional food predator-prey models can cause certain aspects of their dynamics challenging to analyze. In this work, we study a general class of additional food models and establish conditions under which the coexistence equilibrium is globally stable. We then focus on a Holling type IV functional response with AF and show the existence of a Bogdanov-Takens bifurcation of codimension 3. We also study these models through the lens of deterministic chemical reaction network theory. Our analysis shows that the introduction of additional food increases the deficiency of the underlying reaction network and suggests a possible link between higher deficiency and complex bifurcations.

q-bio.PE

Fertilizers Fuel, Insecticides Stabilize: Resolving the Paradox of Enrichment in Agriculture

The Paradox of Enrichment (PoE) predicts that increasing resources, such as nutrient inputs like fertilizers or food availability, should destabilize ecological systems, such as crop-pest dynamics, leading to population cycles that can increase the risk of crop failure during environmental shocks. Yet, since the Green Revolution, fertilizer use has surged without widespread evidence of yield instability, challenging the PoE's relevance to modern agriculture. Here, we propose and test a novel resolution: that insecticides, frequently co-applied with fertilizers, act as stabilizing agents that counterbalance enrichment-induced instability. Using a modified PoE model with empirically grounded parameters for three major crop-pest systems-soybean-aphid, wheat-aphid, and cabbage-diamondback moth-we find that fertilizer increases yields, but destabilizes dynamics, whereas insecticides restore stability and ensure more predictable harvests. These findings reveal that insecticides may suppress pests but also play a critical role in stabilizing crop yields in nutrient-enriched agroecosystems, with implications for ecosystem management, eutrophication, conservation biology, and pesticide policy.

q-bio.PE

An "adaptive" approach to control explosive aphid populations

Classical models of aphid population dynamics are unable to explain multi-peak patterns in field populations. We consider the variable carrying capacity model (VCM), which can generate such complex multi-peak dynamics, but is also demonstrated to show finite-time blow-up behavior via a sign switching structural instability. We build an adaptive behavioral model with a density-dependent switch to stabilize growth, effectively eliminating blow-up, and also capable of generating multiple peaks. Furthermore, guided by empirical work on environment drivers for pests, we devise a non-autonomous model with time-dependent host plant fitness, successfully connecting transient population dynamics with abiotic drivers such as flooding. Finally, we discuss the practical significance of the results through the Economic Threshold (ET) and Economic Injury Level (EIL) calculation for all models. Our simulations all clearly show that aphid abundances exceed these threshold levels, and control is required. Our work provides a stable and biologically relevant prediction scheme for pest outbreaks and their management strategy.

q-bio.PE

An additional food driven biological control patch model, incorporating generalized competition

Additional food sources for an introduced predator are known to increase its efficiency on a target pest. In this context, inhibiting factors such as interference, predator competition, and the introduction of temporally dependent quantity and quality of additional food are all known to enable pest extinction. As climate change and habitat degradation have increasing effects in enhancing patchiness in ecological systems, the effect of additional food in patch models has also been recently considered. However, the question of complete pest extinction in such patchy systems remains open. In the current manuscript, we consider a biological control model where additional food drives competition among predators in one patch, and they subsequently disperse to a neighboring patch via drift or dispersal. We show that complete pest extinction in both patches is possible. Further, this state is proved to be globally asymptotically stable under certain parametric restrictions. We also prove a codimension-2 Bogdanov-Takens bifurcation. We discuss our results in the context of designing pest management strategies under enhanced climate change and habitat fragmentation. Such strategies are particularly relevant to control invasive pests such as the Soybean aphid (\emph{Aphis glycines}), in the North Central United States.

q-bio.PE

Invasive species control via a discrete model for the Trojan Y-chromosome strategy

Invasive species are a growing threat to ecosystems, particularly in aquatic environments. The Trojan Y Chromosome (TYC) strategy is a promising biological method for reducing invasive populations by introducing genetically modified males (supermales) that produce only male offspring, leading to population decline due to a shortage of females. In this study, we develop a novel discrete--time, age--structured mathematical model to simulate the effects of this strategy. Our model divides the life cycle of species into two stages--egg and maturity--and tracks different sub--populations, including supermales. We analyze the equilibria of the system and prove the existence and stability of extinction and positive equilibrium points. Numerical simulations show that extinction depends on factors such as fecundity, the number of supermales released, and initial population sizes. The model also reveals complex behaviors, such as bistability and thresholds for population collapse. This discrete approach offers a useful framework for understanding and optimizing the TYC strategy and can help guide future field applications of invasive species control.

q-bio.PE

Towards improved pest management of the soybean aphid

The soybean aphid (\emph{Aphis glycines}) is an invasive insect pest that continues to cause large-scale damage to soybean crops in the North Central United States. The current manuscript proposes several mathematical models for the top-down bio-control of the aphid, as well as control via pesticides and neonicotinoids. The models are motivated empirically, and constructed based on laboratory experiments conducted to test control of aphids by Lacewing larvae, as well as by a parasitic wasp (\emph{Aphidius colemani}). The effectiveness of these models is compared by taking into account factors such as economic injury levels for soybeans, life history traits such as cannibalism amongst the predator, and intraguild predation between competing bio-control agents such as predators and parasitoids. The models predict multiple population peaks and transient chaotic dynamics when a predator and/or insecticides are used. It is observed that parasitoids, in conjunction with predators, are more efficient at stabilizing the population dynamics than insecticide use. They also suggest a combination of predators, parasitoids, and insecticides would be more efficient at suppressing aphid populations than using only predators or parasitoids. The models also qualitatively capture the features seen in long-time field data from 2000-2013. We discuss applications of our results to pest management strategies for soybean aphids in the context of a changing climate, as well as regime shifts.

q-bio.PE

An eco-epidemiological model with prey-taxis and slow diffusion: Global existence, boundedness and novel dynamics

In this manuscript, an attempt has been made to understand the effects of prey-taxis on the existence of global-in-time solutions and dynamics in an eco-epidemiological model, particularly under the influence of slow dispersal characterized by the $p$-Laplacian operator and enhanced mortality of the infected prey, subject to specific assumptions on the taxis sensitivity functions. We prove the global existence of classical solutions when the infected prey undergoes random motion and exhibits standard mortality. Under the assumption that the infected prey disperses slowly and exhibits enhanced mortality, we prove the global existence of weak solutions. Following a detailed mathematical investigation of the proposed model, we shift our focus to analyse the stability of the positive equilibrium point under the scenario where all species exhibit linear diffusion, the infected prey experiences standard mortality, and the predator exhibits taxis exclusively toward the infected prey. Within this framework, we establish the occurrence of a steady-state bifurcation. Numerical simulations are then carried out to observe this dynamical behavior. Our results have large scale applications to biological invasions and biological control of pests, under the prevalence of disease in the pest population.

math.AP

Existence of periodic solution of a non-autonomous allelopathic phytoplankton model with fear effect

In this paper, we consider a non-autonomous allelopathic phytoplankton competition ODE model, incorporating the influence of fear effects observed in natural biological phenomena. Based on Mawhin's coincidence degree theory some sufficient conditions for existence of periodic solutions are obtained. We validate our findings through an illustrative example and numerical simulations, showing that constant coefficients lead to steady-state dynamics, while periodic variations induce oscillatory behavior.

math.DS

T(w)o patch or not t(w)o patch: A novel additional food model

A number of top down bio-control models have been proposed where the introduced predators' efficacy is enhanced via the provision of additional food (AF). However, if the predator has a pest dependent monotone functional response, pest extinction is unattainable. In the current manuscript, we propose a model where a predator with pest dependent monotone functional response is introduced into a ``patch" such as a prairie strip with AF, and then disperses or drifts into a neighboring ``patch" such as a crop field, to target a pest. We show the pest extinction state is attainable in the crop field and can be globally attracting. The AF model with patch structure can eliminate predator explosion present therein and can keep pest densities lower than the classical top-down bio-control model. We provide the first proof of the global stability of the interior equilibrium for the classical AF model. We also observe ``patch-specific chaos" - the pest occupying the crop field can oscillate chaotically, while the pest in the prairie strip oscillates periodically. We discuss these results in light of bio-control strategies that utilize state-of-the-art farming practices such as prairie strips and drift and dispersal pressures driven by climate change.

q-bio.PE

Novel Dynamics in an Additional Food provided Predator-Prey System with mutual interference

The provision of additional food (AF) sources to an introduced predator has been identified as a mechanism to improve pest control. However, AF models with prey dependent functional responses can cause unbounded growth of the predator \cite{S27}. To avoid such dynamics, an AF model with mutual interference effect has been proposed \cite{S02}. The analysis therein reveals that if the quantity of additional food $ξ> h(ε)$, where $ε$ is the mutual interference parameter, then pest eradication is possible, and this is facilitated via a transcritical bifurcation. We revisit this model and show novel dynamical behaviors. In particular, pest eradication is possible for a tighter range of AF $g(ε) < ξ< f(ε) < h(ε)$, and can also occur via a saddle node bifurcation. We observe bi-stability, as well as local bifurcations of Hopf type. We also prove a global bifurcation, of homoclinic type. This bifurcation in turn is shown to create a non-standard dynamic wherein the pest extinction state becomes an ``almost" global attractor. To the best of our knowledge, this is the first proof of existence of such a dynamical structure in AF models. We discuss our analysis in the context of designing novel bio-control strategies.

math.DS

Stability and bifurcation analysis of a two-patch model with Allee effect and dispersal

In the current manuscript, a first two-patch model with Allee effect and nonlinear dispersal is presented. We study both the ODE case and the PDE case here. In the ODE model, the stability of the equilibrium points and the existence of saddle-node bifurcation are discussed. The phase diagram and bifurcation curve of our model are also given by numerical simulation. Besides, the corresponding linear dispersal case is also presented. We show that when the Allee effect is large, high intensity of linear dispersal is not favorable to the persistence of the species. We further show when the Allee effect is large, nonlinear diffusion is more favorable to the survival of the population than linear diffusion. Moreover, the results of the PDE model extends our findings from discrete patches to continuous patches.

math.DS

Dynamical Analysis of an Allelopathic Phytoplankton Model with Fear Effect

This paper is the first to propose an allelopathic phytoplankton competition ODE model influenced by a fear effect based on natural biological phenomena. It is shown that the interplay of this fear effect and the allelopathic term cause rich dynamics in the proposed competition model, such as global stability, transcritical bifurcation, pitchfork bifurcation, and saddle-node bifurcation. We also consider the spatially explicit version of the model and prove analogous results. Numerical simulations verify the feasibility of the theoretical analysis. The results demonstrate that the primary cause of the extinction of non-toxic species is the fear of toxic species compared to toxins. Allelopathy only affects the density of non-toxic species. The discussion provides guidance for the conservation of species and the maintenance of biodiversity.

math.DS

Exploring unique dynamics in a predator-prey model with generalist predator and group defence in prey

In the current manuscript, we consider a predator-prey model where the predator is modeled as a generalist using a modified Leslie-Gower scheme, and the prey exhibits group defence via a generalised response. We show that the model could exhibit finite time blow-up, contrary to the current literature (Eur. Phys. J. Plus 137, 28). We also propose a new concept via which the predator population blows up in finite time while the prey population quenches in finite time. The blow-up and quenching times are proved to be one and the same. Our analysis is complemented by numerical findings. This includes a numerical description of the basin of attraction for large data blow-up solutions, as well as several rich bifurcations leading to multiple limit cycles, both in co-dimension one and two. Lastly, we posit a delayed version of the model with globally existing solutions for any initial data.

math.DS

The effect of "very fast" strategies on two species competition

We consider the effect of finite time extinction mechanisms (FTEM) such as (1) semi-linear harvesting terms, and (2) quasi-linear fast diffusion terms on two species Lokta-Volterra competition models. We show that these mechanisms can alter classical dynamics of competitive exclusion, and weak and strong competition by acting only on a \emph{small} portion of the weaker competitors' population, analogous to small defector populations in game theory \cite{DC23}. In particular, a stronger competitors population, with a few individuals dispersing (``defecting") very quickly, could exhibit bi-stability, as well as competitive exclusion \emph{reversal}. The non-linear harvesting is applied to aphid-soybean crop systems, wherein novel dynamics are observed. Applications to bio-control of invasive pests such as the soybean aphid are discussed.

math.DS

Dynamical Analysis of a Lotka-Volterra Competition Model with both Allee and Fear Effect

Population ecology theory is replete with density dependent processes. However trait-mediated or behavioral indirect interactions can both reinforce or oppose density-dependent effects. This paper presents the first two species competitive ODE and PDE systems where an Allee effect, which is a density dependent process and the fear effect, which is non-consumptive and behavioral are both present. The stability of the equilibria is discussed analytically using the qualitative theory of ordinary differential equations. It is found that the Allee effect and the fear effect change the extinction dynamics of the system and the number of positive equilibrium points, but they do not affect the stability of the positive equilibria. We also observe some special dynamics that induce bifurcations in the system by varying the Allee or fear parameter. Interestingly we find that the Allee effect working in conjunction with the fear effect, can bring about several qualitative changes to the dynamical behavior of the system with only the fear effect in place, in regimes of small fear. That is, for small amounts of the fear parameter, it can change a competitive exclusion type situation to a strong competition type situation. It can also change a weak competition type situation to a bi-stability type situation. However for large fear regimes the Allee effect reinforces the dynamics driven by the fear effect. The analysis of the corresponding spatially explicit model is also presented. To this end the comparison principle for parabolic PDE is used. The conclusions of this paper have strong implications for conservation biology, biological control as well as the preservation of biodiversity.

q-bio.PE