arXiv subjects

Sylvie Méléard

Publications and source records attributed to Sylvie Méléard.

49 records · Page 3Linked to original sources

arXiv2011

Slow and fast scales for superprocess limits of age-structured populations

A superprocess limit for an interacting birth-death particle system modelling a population with trait and physical age-structures is established. Traits of newborn offspring are inherited from the parents except when mutations occur, while ages are set to zero. Because of interactions between individuals, standard approaches based on the Laplace transform do not hold. We use a martingale problem approach and a separation of the slow (trait) and fast (age) scales. While the trait marginals converge in a pathwise sense to a superprocess, the age distributions, on another time scale, average to equilibria that depend on traits. The convergence of the whole process depending on trait and age, only holds for finite-dimensional time-marginals. We apply our results to the study of examples illustrating different cases of trade-off between competition and senescence.

math.PR↗

arXiv2011

Lévy flights in ecology

We are interested in modeling Darwinian evolution resulting from the interplay of phenotypic variation and natural selection through ecological interactions. The population is modeled as a stochastic point process whose generator captures the probabilistic dynamics over continuous time of birth, mutation, and death, as influenced by each individual's trait values, and interactions between individuals. An offspring usually inherits the trait values of her progenitor, except when a random mutation causes the offspring to take an instantaneous mutation step at birth to new trait values. In the case we are interested in, the probability distribution of mutations has a heavy tail and belongs to the domain of attraction of a stable law. We investigate the large-population limit with allometric demographies: larger populations made up of smaller individuals which reproduce and die faster, as is typical for micro-organisms. We show that depending on the allometry coefficient the limit behavior of the population process can be approximated by nonlinear Lévy flights of different nature: either deterministic, in the form of nonlocal fractional reaction-diffusion equations, or stochastic, as nonlinear super-processes with the underlying reaction and a fractional diffusion operator. These approximation results demonstrate the existence of such nontrivial fractional objects; their uniqueness is also proved.

math.PR↗

arXiv2011

A host-parasite multilevel interacting process and continuous approximations

We are interested in modeling some two-level population dynamics, resulting from the interplay of ecological interactions and phenotypic variation of individuals (or hosts) and the evolution of cells (or parasites) of two types living in these individuals. The ecological parameters of the individual dynamics depend on the number of cells of each type contained by the individual and the cell dynamics depends on the trait of the invaded individual. Our models are rooted in the microscopic description of a random (discrete) population of individuals characterized by one or several adaptive traits and cells characterized by their type. The population is modeled as a stochastic point process whose generator captures the probabilistic dynamics over continuous time of birth, mutation and death for individuals and birth and death for cells. The interaction between individuals (resp. between cells) is described by a competition between individual traits (resp. between cell types). We look for tractable large population approximations. By combining various scalings on population size, birth and death rates and mutation step, the single microscopic model is shown to lead to contrasting nonlinear macroscopic limits of different nature: deterministic approximations, in the form of ordinary, integro- or partial differential equations, or probabilistic ones, like stochastic partial differential equations or superprocesses. The study of the long time behavior of these processes seems very hard and we only develop some simple cases enlightening the difficulties involved.

math.PR↗

arXiv2010

Uniform estimates for metastable transition times in a coupled bistable system

We consider a coupled bistable N-particle system driven by a Brownian noise, with a strong coupling corresponding to the synchronised regime. Our aim is to obtain sharp estimates on the metastable transition times between the two stable states, both for fixed N and in the limit when N tends to infinity, with error estimates uniform in N. These estimates are a main step towards a rigorous understanding of the metastable behavior of infinite dimensional systems, such as the stochastically perturbed Ginzburg-Landau equation. Our results are based on the potential theoretic approach to metastability.

math.PR↗

arXiv2009

Quasi-stationary distributions for structured birth and death processes with mutations

We study the probabilistic evolution of a birth and death continuous time measure-valued process with mutations and ecological interactions. The individuals are characterized by (phenotypic) traits that take values in a compact metric space. Each individual can die or generate a new individual. The birth and death rates may depend on the environment through the action of the whole population. The offspring can have the same trait or can mutate to a randomly distributed trait. We assume that the population will be extinct almost surely. Our goal is the study, in this infinite dimensional framework, of quasi-stationary distributions when the process is conditioned on non-extinction. We firstly show in this general setting, the existence of quasi-stationary distributions. This result is based on an abstract theorem proving the existence of finite eigenmeasures for some positive operators. We then consider a population with constant birth and death rates per individual and prove that there exists a unique quasi-stationary distribution with maximal exponential decay rate. The proof of uniqueness is based on an absolute continuity property with respect to a reference measure.

math.PR↗

arXiv2009

Quasi-stationary distributions and diffusion models in population dynamics

In this paper, we study quasi-stationarity for a large class of Kolmogorov diffusions. The main novelty here is that we allow the drift to go to $- \infty$ at the origin, and the diffusion to have an entrance boundary at $+\infty$. These diffusions arise as images, by a deterministic map, of generalized Feller diffusions, which themselves are obtained as limits of rescaled birth--death processes. Generalized Feller diffusions take nonnegative values and are absorbed at zero in finite time with probability 1. An important example is the logistic Feller diffusion. We give sufficient conditions on the drift near 0 and near $+ \infty$ for the existence of quasi-stationary distributions, as well as rate of convergence in the Yaglom limit and existence of the $Q$-process. We also show that under these conditions, there is exactly one quasi-stationary distribution, and that this distribution attracts all initial distributions under the conditional evolution, if and only if $+\infty$ is an entrance boundary. In particular this gives a sufficient condition for the uniqueness of quasi-stationary distributions. In the proofs spectral theory plays an important role on $L^2$ of the reference measure for the killed process.

math.PR↗

arXiv2008

Polymorphic evolution sequence and evolutionary branching

We are interested in the study of models describing the evolution of a polymorphic population with mutation and selection in the specific scales of the biological framework of adaptive dynamics. The population size is assumed to be large and the mutation rate small. We prove that under a good combination of these two scales, the population process is approximated in the long time scale of mutations by a Markov pure jump process describing the successive trait equilibria of the population. This process, which generalizes the so-called trait substitution sequence, is called polymorphic evolution sequence. Then we introduce a scaling of the size of mutations and we study the polymorphic evolution sequence in the limit of small mutations. From this study in the neighborhood of evolutionary singularities, we obtain a full mathematical justification of a heuristic criterion for the phenomenon of evolutionary branching. To this end we finely analyze the asymptotic behavior of 3-dimensional competitive Lotka-Volterra systems.

math.PR↗

arXiv2008

Competitive or Weak Cooperative Stochastic Lotka-Volterra Systems Conditioned to Non-Extinction

We are interested in the long time behavior of a two-type density-dependent biological population conditioned to non-extinction, in both cases of competition or weak cooperation between the two species. This population is described by a stochastic Lotka-Volterra system, obtained as limit of renormalized interacting birth and death processes. The weak cooperation assumption allows the system not to blow up. We study the existence and uniqueness of a quasi-stationary distribution, that is convergence to equilibrium conditioned to non extinction. To this aim we generalize in two-dimensions spectral tools developed for one-dimensional generalized Feller diffusion processes. The existence proof of a quasi-stationary distribution is reduced to the one for a $d$-dimensional Kolmogorov diffusion process under a symmetry assumption. The symmetry we need is satisfied under a local balance condition relying the ecological rates. A novelty is the outlined relation between the uniqueness of the quasi-stationary distribution and the ultracontractivity of the killed semi-group. By a comparison between the killing rates for the populations of each type and the one of the global population, we show that the quasi-stationary distribution can be either supported by individuals of one (the strongest one) type or supported by individuals of the two types. We thus highlight two different long time behaviors depending on the parameters of the model: either the model exhibits an intermediary time scale for which only one type (the dominant trait) is surviving, or there is a positive probability to have coexistence of the two species.

math.PR↗

arXiv2007

Age-structured Trait Substitution Sequence Process and Canonical Equation

We are interested in a stochastic model of trait and age-structured population undergoing mutation and selection. We start with a continuous time, discrete individual-centered population process. Taking the large population and rare mutations limits under a well-chosen time-scale separation condition, we obtain a jump process that generalizes the Trait Substitution Sequence process describing Adaptive Dynamics for populations without age structure. Under the additional assumption of small mutations, we derive an age-dependent ordinary differential equation that extends the Canonical Equation. These evolutionary approximations have never been introduced to our knowledge. They are based on ecological phenomena represented by PDEs that generalize the Gurtin-McCamy equation in Demography. Another particularity is that they involve a fitness function, describing the probability of invasion of the resident population by the mutant one, that can not always be computed explicitly. Examples illustrate how adding an age-structure enrich the modelling of structured population by including life history features such as senescence. In the cases considered, we establish the evolutionary approximations and study their long time behavior and the nature of their evolutionary singularities when computation is tractable. Numerical procedures and simulations are carried.

math.PR↗

arXiv2007

Nonlinear SDEs driven by Lévy processes and related PDEs

In this paper we study general nonlinear stochastic differential equations, where the usual Brownian motion is replaced by a Lévy process. We also suppose that the coefficient multiplying the increments of this process is merely Lipschitz continuous and not necessarily linear in the time-marginals of the solution as is the case in the classical McKean-Vlasov model. We first study existence, uniqueness and particle approximations for these stochastic differential equations. When the driving process is a pure jump Lévy process with a smooth but unbounded Lévy measure, we develop a stochastic calculus of variations to prove that the time-marginals of the solutions are absolutely continuous with respect to the Lebesgue measure. In the case of a symmetric stable driving process, we deduce the existence of a function solution to a nonlinear integro-differential equation involving the fractional Laplacian.

math.PR↗

arXiv2006

Invasion and adaptive evolution for individual-based spatially structured populations

The interplay between space and evolution is an important issue in population dynamics, that is in particular crucial in the emergence of polymorphism and spatial patterns. Recently, biological studies suggest that invasion and evolution are closely related. Here we model the interplay between space and evolution starting with an individual-based approach and show the important role of parameter scalings on clustering and invasion. We consider a stochastic discrete model with birth, death, competition, mutation and spatial diffusion, where all the parameters may depend both on the position and on the trait of individuals. The spatial motion is driven by a reflected diffusion in a bounded domain. The interaction is modelled as a trait competition between individuals within a given spatial interaction range. First, we give an algorithmic construction of the process. Next, we obtain large population approximations, as weak solutions of nonlinear reaction-diffusion equations with Neumann's boundary conditions. As the spatial interaction range is fixed, the nonlinearity is nonlocal. Then, we make the interaction range decrease to zero and prove the convergence to spatially localized nonlinear reaction-diffusion equations, with Neumann's boundary conditions. Finally, simulations based on the microscopic individual-based model are given, illustrating the strong effects of the spatial interaction range on the emergence of spatial and phenotypic diversity (clustering and polymorphism) and on the interplay between invasion and evolution. The simulations focus on the qualitative differences between local and nonlocal interactions.

math.PR↗

arXiv2005

Individual-based probabilistic models of adaptive evolution and various scaling approximations

We are interested in modelling Darwinian evolution, resulting from the interplay of phenotypic variation and natural selection through ecological interactions. Our models are rooted in the microscopic, stochastic description of a population of discrete individuals characterized by one or several adaptive traits. The population is modelled as a stochastic point process whose generator captures the probabilistic dynamics over continuous time of birth, mutation, and death, as influenced by each individual's trait values, and interactions between individuals. An offspring usually inherits the trait values of her progenitor, except when a mutation causes the offspring to take an instantaneous mutation step at birth to new trait values. We look for tractable large population approximations. By combining various scalings on population size, birth and death rates, mutation rate, mutation step, or time, a single microscopic model is shown to lead to contrasting macroscopic limits, of different nature: deterministic, in the form of ordinary, integro-, or partial differential equations, or probabilistic, like stochastic partial differential equations or superprocesses. In the limit of rare mutations, we show that a possible approximation is a jump process, justifying rigorously the so-called trait substitution sequence. We thus unify different points of view concerning mutation-selection evolutionary models.

math.PR↗

arXiv2005

Estimates for the density of a nonlinear Landau process

The aim of this paper is to obtain estimates for the density of the law of a specific nonlinear diffusion process at any positive bounded time. This process is issued from kinetic theory and is called Landau process, by analogy with the associated deterministic Fokker-Planck-Landau equation. It is not Markovian, its coefficients are not bounded and the diffusion matrix is degenerate. Nevertheless, the specific form of the diffusion matrix and the nonlinearity imply the non-degeneracy of the Malliavin matrix and then the existence and smoothness of the density. In order to obtain a lower bound for the density, the known results do not apply. However, our approach follows the main idea consisting in discretizing the interval time and developing a recursive method. To this aim, we prove and use refined results on conditional Malliavin calculus. The lower bound implies the positivity of the solution of the Landau equation, and partially answers to an analytical conjecture. We also obtain an upper bound for the density, which again leads to an unusual estimate due to the bad behavior of the coefficients.

math.PR↗