Kernel-Dependent Pattern Formation in a Population Model with Nonlocal Facilitation and Competition

Abstract

Spatial patterns, such as those in dryland vegetation models, have historically been studied in systems of reaction-diffusion systems with pattern onset via a Turing bifurcation from a spatially uniform state. More recently, spatial patterns have been considered in models that incorporate spatially extended interactions via nonlocal interaction kernels. It remains largely underexplored if and how the choice of nonlocal interaction kernel contributes to differences in pattern formation and persistence, particularly in models that contain competition and facilitation. Here, we investigate spatial patterns in a reaction-diffusion model for a single species that includes nonlocal competition and facilitation processes; Gaussian, exponential, algebraic, hat, and smooth hat kernels are considered as specific examples. Via a center manifold analysis, and using the relative spatial scale of competition to facilitation and the death rate as bifurcation parameters, we identify that the choice of kernel has impacts on the pattern forming bifurcation. Bifurcations using the Gaussian, exponential, and algebraic kernels largely follow expectations of Turing patterns, but patterns in the hat and smooth hat kernels can form even when the scale of competition is less than that of facilitation. The dynamics of patterns far from onset are investigated via numerical continuation methods. The model produces the so-called "Turing-before-Tipping" phenomenon demonstrating that the arrangement into spatial patterns is an effective resilience mechanism against harsh conditions. Again, there is a kernel-dependent dichotomy in pattern behavior. Early warning signs for population extinction are observed with the Gaussian, exponential, and algebraic kernels, but not under the hat or smooth hat cases.