SearcharxivSearch

arXiv subjects

Alan McKane

Publications and source records attributed to Alan McKane.

12 recordsLinked to original sources

Enhanced stochastic oscillations in autocatalytic reactions

We study a simplified scheme of $k$ coupled autocatalytic reactions, previously introduced by Togashi and Kaneko. The role of stochastic fluctuations is elucidated through the use of the van Kampen system-size expansion and the results compared with direct stochastic simulations. Regular temporal oscillations are predicted to occur for the concentration of the various chemical constituents, with an enhanced amplitude resulting from a resonance which is induced by the intrinsic graininess of the system. The associated power spectra are determined and have a different form depending on the number of chemical constituents, $k$. We make detailed comparisons in the two cases $k=4$ and $k=8$. Agreement between the theoretical and numerical results for the power spectrum are good in both cases. The resulting spectrum is especially interesting in the $k=8$ system, since it has two peaks, which the system-size expansion is still able to reproduce accurately.

cond-mat.stat-mech

Topological structure and interaction strengths in model food webs

We report the results of carrying out a large number of simulations on a coevolutionary model of multispecies communities. A wide range of parameter values were investigated which allowed a rather complete picture of the change in behaviour of the model as these parameters were varied to be built up. Our main interest was in the nature of the community food webs constructed via the simulations. We identify the range of parameter values which give rise to realistic food webs and give arguments which allow some of the structure which is found to be understood in an intuitive way. Since the webs are evolved according to the rules of the model, the strengths of the predator-prey links are not determined a priori, and emerge from the process of constructing the web. We measure the distribution of these link strengths, and find that there are a large number of weak links, in agreement with recent suggestions. We also review some of the data on food webs available in the literature, and make some tentative comparisons with our results. The difficulties of making such comparisons and the possible future developments of the model are also briefly discussed.

q-bio.PE

Deleting species from model food webs

We use food webs generated by a model to investigate the effects of deleting species on other species in the web and on the web as a whole. The model incorporates a realistic population dynamics, adaptive foragers and other features which allow for the construction of model webs which resemble empirical food webs. A large number of simulations were carried out to produce a substantial number of model webs on which deletion experiments could be performed. We deleted each species in four hundred distinct model webs and determined, on average, how many species were eliminated from the web as a result. Typically only a small number of species became extinct; in no instance was the web close to collapse. Next, we examined how the the probability of extinction of a species depended on its relationship with the deleted species. This involved the exploration of the concept of indirect predator and prey species and the extent that the probability of extinction depended on the trophic level of the two species. The effect of deletions on the web itself was studied by searching for keystone species, whose removal caused a major restructuring of the community, and also by looking at the correlation between a number of food web properties (number of species, linkage density, fraction of omnivores, degree of cycling and redundancy) and the stability of the web to deletions. With the exception of redundancy, we found little or no correlation. In particular, we found no evidence that complexity in terms of increased species number or links per species is destabilising.

q-bio.PE

The impact of non-linear functional responses on the long-term evolution of food web structure

We investigate the long-term web structure emerging in evolutionary food web models when different types of functional responses are used. We find that large and complex webs with several trophic layers arise only if the population dynamics is such that it allows predators to focus on their best prey species. This can be achieved using modified Lotka-Volterra or Holling/Beddington functional responses with effective couplings that depend on the predator's efficiency at exploiting the prey, or a ratio-dependent functional response with adaptive foraging. In contrast, if standard Lotka-Volterra or Holling/Beddington functional responses are used, long-term evolution generates webs with almost all species being basal, and with additionally many links between these species. Interestingly, in all cases studied, a large proportion of weak links result naturally from the evolution of the food webs.

q-bio.PE

Analytic solution of Hubbell's model of local community dynamics

Recent theoretical approaches to community structure and dynamics reveal that many large-scale features of community structure (such as species-rank distributions and species-area relations) can be explained by a so-called neutral model. Using this approach, species are taken to be equivalent and trophic relations are not taken into account explicitly. Here we provide a general analytic solution to the local community model of Hubbell's neutral theory of biodiversity by recasting it as an urn model i.e.a Markovian description of states and their transitions. Both stationary and time-dependent distributions are analysed. The stationary distribution -- also called the zero-sum multinomial -- is given in closed form. An approximate form for the time-dependence is obtained by using an expansion of the master equation. The temporal evolution of the approximate distribution is shown to be a good representation for the true temporal evolution for a large range of parameter values.

physics.bio-ph

Functional determinants by contour integration methods

We present a simple and accessible method which uses contour integration methods to derive formulae for functional determinants. To make the presentation as clear as possible, the general idea is first illustrated on the simplest case: a second order differential operator with Dirichlet boundary conditions. The method is applicable to more general situations, and we discuss the way in which the formalism has to be developed to cover these cases. In particular, we also show that simple and elegant formulae exist for the physically important case of determinants where zero modes exist, but have been excluded.

math-ph

Unstable decay and state selection II

The decay of unstable states when several metastable states are available for occupation is investigated using path-integral techniques. Specifically, a method is described which allows the probabilities with which the metastable states are occupied to be calculated by finding optimal paths, and fluctuations about them, in the weak noise limit. The method is illustrated on a system described by two coupled Langevin equations, which are found in the study of instabilities in fluid dynamics and superconductivity. The problem involves a subtle interplay between non-linearities and noise, and a naive approximation scheme which does not take this into account is shown to be unsatisfactory. The use of optimal paths is briefly reviewed and then applied to finding the conditional probability of ending up in one of the metastable states, having begun in the unstable state. There are several aspects of the calculation which distinguish it from most others involving optimal paths: (i) the paths do not begin and end on an attractor, and moreover, the final point is to a large extent arbitrary, (ii) the interplay between the fluctuations and the leading order contribution are at the heart of the method, and (iii) the final result involves quantities which are not exponentially small in the noise strength. This final result, which gives the probability of a particular state being selected in terms of the parameters of the dynamics, is remarkably simple and agrees well with the results of numerical simulations. The method should be applicable to similar problems in a number of other areas such as state selection in lasers, activationless chemical reactions and population dynamics in fluctuating environments.

cond-mat.stat-mech

Renormalization group and perfect operators for stochastic differential equations

We develop renormalization group methods for solving partial and stochastic differential equations on coarse meshes. Renormalization group transformations are used to calculate the precise effect of small scale dynamics on the dynamics at the mesh size. The fixed point of these transformations yields a perfect operator: an exact representation of physical observables on the mesh scale with minimal lattice artifacts. We apply the formalism to simple nonlinear models of critical dynamics, and show how the method leads to an improvement in the computational performance of Monte Carlo methods.

cond-mat.stat-mech

A mean field stochastic theory for species-rich assembled communities

A dynamical model of an ecological community is analyzed within a "mean-field approximation" in which one of the species interacts with the combination of all of the other species in the community. Within this approximation the model may be formulated as a master equation describing a one-step stochastic process. The stationary distribution is obtained in closed form and is shown to reduce to a logseries or lognormal distribution, depending on the values that the parameters describing the model take on. A hyperbolic relationship between the connectance of the matrix of interspecies interactions and the average number of species, exists for a range of parameter values. The time evolution of the model at short and intermediate times is analyzed using van Kampen's approximation, which is valid when the number of individuals in the community is large. Good agreement with numerical simulations is found. The large time behavior, and the approach to the stationary state, is obtained by solving the equation for the generating function of the probability distribution. The analytical results which follow from the analysis are also in good agreement with direct simulations of the model.

nlin.AO

Integrated random processes exhibiting long tails, finite moments and 1/f spectra

A dynamical model based on a continuous addition of colored shot noises is presented. The resulting process is colored and non-Gaussian. A general expression for the characteristic function of the process is obtained, which, after a scaling assumption, takes on a form that is the basis of the results derived in the rest of the paper. One of these is an expansion for the cumulants, which are all finite, subject to mild conditions on the functions defining the process. This is in contrast with the Levy distribution -which can be obtained from our model in certain limits- which has no finite moments. The evaluation of the power spectrum and the form of the probability density function in the tails of the distribution shows that the model exhibits a 1/f spectrum and long tails in a natural way. A careful analysis of the characteristic function shows that it may be separated into a part representing a Levy processes together with another part representing the deviation of our model from the Levy process. This allows our process to be viewed as a generalization of the Levy process which has finite moments.

cond-mat.stat-mech

Scaling in a Multispecies Network Model Ecosystem

A new model ecosystem consisting of many interacting species is introduced. The species are connected through a random matrix with a given connectivity. It is shown that the system is organized close to a boundary of marginal stability in such a way that fluctuations follow power law distributions both in species abundance and their lifetimes for some slow-driving (immigration) regime. The connectivity and the number of species are linked through a scaling relation which is the one observed in real ecosystems. These results suggest that the basic macroscopic features of real, species-rich ecologies might be linked with a critical state. A natural link between lognormal and power law distributions of species abundances is suggested.

adap-org

Block Spins for Partial Differential Equations

We investigate the use of renormalisation group methods to solve partial differential equations (PDEs) numerically. Our approach focuses on coarse-graining the underlying continuum process as opposed to the conventional numerical analysis method of sampling it. We calculate exactly the coarse-grained or `perfect' Laplacian operator and investigate the numerical effectiveness of the technique on a series of 1+1-dimensional PDEs with varying levels of smoothness in the dynamics: the diffusion equation, the time-dependent Ginzburg-Landau equation, the Swift-Hohenberg equation and the damped Kuramoto-Sivashinsky equation. We find that the renormalisation group is superior to conventional sampling-based discretisations in representing faithfully the dynamics with a large grid spacing, introducing no detectable lattice artifacts as long as there is a natural ultra-violet cut off in the problem. We discuss limitations and open problems of this approach.

cond-mat.stat-mech