arXiv subjects

Mehran Kardar

Publications and source records attributed to Mehran Kardar.

At least 19 recordsLinked to original sources

arXiv2026

Heat Transfer and Torque in Enclosing Cylindrical Configurations with Nonreciprocal Materials

Electromagnetic fluctuations can transfer not only energy but also angular momentum, leading to forces, torques, heat currents, and friction in out-of-equilibrium setups. In enclosing configurations, we show that if at least one of two objects is rotationally symmetric, the torque is bounded by heat transfer, since both arise from photon transfers with angular momentum $\hbar n$ and energy $\hbar\omega$. With only one object assumed to be rotationally symmetric, it may be possible to obtain a nonzero torque with reciprocal media, but nonreciprocal media are required to break the symmetry between $n$ and $-n$ and produce a nonzero torque if both objects are rotationally symmetric. We then specialize to concentric cylinders with a nonreciprocal dielectric response and use Rytov fluctuational electrodynamics to express heat transfer and torque in terms of an angular-momentum-resolved flux density, $\Phi_n(\omega)$. We also analyze the conditions for stable levitation of the inner cylinder using the proximity force approximation, in the process obtaining a new analytic formula for the normal Casimir force between dilute plates at different temperatures. Finally, to find the extracted work in a contactless engine setup, we compute the fluctuation-induced friction for a slowly rotating inner cylinder, and we find a bound between torque, friction, and heat transfer. Due to this bound, the efficiency of the heat engine remains bounded by the Carnot limit.

cond-mat.stat-mech↗

arXiv2026

A Spectral Route to Directed-Polymer Glasses

A finite density of mutually avoiding directed polymers in a quenched random medium is a minimal model of glassy line matter. The dilute theory, solved by replica Bethe ansatz, predicts an interaction free energy proportional to $\rho^2$ and disorder cumulants with distinct power-law dependences on the density $\rho$, but direct numerical tests have been hindered by the combinatorially large many-polymer transfer matrix. We recast the problem as filling logarithmic eigenvalues of a single-polymer transfer-matrix product, obtaining the quenched free energy, its cumulants, and a disorder-induced linear spectral edge consistent with the replica prediction.

cond-mat.dis-nn↗

arXiv2026

A Contactless Heat Engine Driven by Nonreciprocal Fluctuation-Induced Torques

We describe a contactless heat engine in which quantum and thermal electromagnetic fluctuations act as the working medium. The setup consists of two concentric cylinders held at different temperatures. The inner cylinder stably levitates within the outer one due to repulsive nonequilibrium Casimir forces. The chirality of the setup is broken by using nonreciprocal dielectric materials, akin to application of a magnetic field along the common cylinder axis. Using Rytov fluctuational electrodynamics, we show that heat transfer and torque can be expressed in terms of an angular-momentum-resolved heat flux density, $\Phi_n(\omega)$: each exchanged photon carries energy $\hbar \omega$ and angular momentum $\hbar n$. In reciprocal media contributions from modes $n$ and $-n$ cancel and there is no net torque; nonreciprocity breaks this symmetry and powers rotation of the inner cylinder. Even in the absence of contact, electromagnetic fluctuations produce a frictional torque opposing rotation that we compute. This enables computation of characteristic steady state rotations, and estimation of the engine efficiency (which remains bounded by the Carnot limit). The cylindrical setup provides a natural realization of fluctuation-induced angular-momentum transfer and a possible route toward nanoscale contactless engines.

quant-ph↗

arXiv2026

Generic long-range correlations in nonequilibrium mixtures

We study correlation functions in generic non-equilibrium mixtures, including multi-temperature systems and non-reciprocal field theories. The corresponding linear theory is short-ranged, and nonlinearities are irrelevant in the renormalization-group sense. Nonetheless, we find that these nonlinearities generate long-ranged three-point correlations in the isotropic disordered phase. Our analytical predictions, which are based on a phenomenological theory, are confirmed by numerical simulations of Brownian colloids in contact with thermal baths at different temperatures. Dangerously irrelevant nonlinearities in non-equilibrium mixtures thus offer a new route to long-range correlations, supporting the hypothesis that such correlations are not the exception but the rule out of equilibrium.

cond-mat.stat-mech↗

arXiv2026

Competition at the front of expanding populations

When competing species grow into new territory, the population is dominated by descendants of successful ancestors at the expansion front. Successful ancestry depends on both the reproductive advantage (fitness), as well as ability and opportunity to colonize new domains. We present a model that integrates both elements by coupling the classic description of one-dimensional competition (Fisher equation) to the minimal model of front shape (KPZ equation). Macroscopic manifestations of these equations are distinct growth morphologies controlled by expansion rates, competitive abilities, or spatial anisotropy. In some cases the ability to expand in space may overcome reproductive advantage in colonizing new territory. When new traits appear with accumulating mutations, we find that variations in fitness in range expansion may be described by the Tracy--Widom distribution.

q-bio.PE↗

arXiv2026

Directed Polymer Transfer Matrices as a Unified Generator of Distinct One-Point Fluctuation Laws

We numerically revisit the transfer-matrix formulation of directed polymers in random media and show that a common finite-dimensional framework organizes the canonical one-point fluctuation laws in $(1+1)$ dimensions. For a fixed realization of the bulk disorder, full-space partition functions are obtained from the same time-ordered product $W(t)$ through endpoint contractions or a Brownian-weighted initial vector, while the half-space construction modifies only the transfer rule at the absorbing boundary. These choices yield distributions consistent with the standard KPZ subclasses: Tracy--Widom GUE for point-to-point geometry, Tracy--Widom GOE for point-to-line geometry, Tracy--Widom GSE for half-space point-to-point geometry, and Baik--Rains for the stationary line-to-point construction. In all four cases, the free-energy fluctuations grow as $t^{1/3}$, and the low-order cumulants approach the corresponding universal benchmarks. The matrix-product formulation also provides access to intrinsic spectral observables. For the leading eigenvalue $\lambda_1(t)$, the fluctuations of $\ln\lambda_1(t)$ exhibit an intermediate $t^{1/3}$ regime, while the standardized distribution remains distinct from the canonical benchmark laws over the studied time range.

cond-mat.soft↗

arXiv2026

Spatiotemporal noise stabilizes unbounded diversity in strongly-competitive communities

Classical ecological models predict that diverse communities should be unstable, presenting a central challenge to explaining the stable biodiversity seen in nature. We revisit this long-standing problem by extending the generalized Lotka-Volterra model to include both spatial structure and environmental fluctuations across space and time. We find that neither space nor environmental noise alone can resolve the tension between diversity and stability, but that together they permit arbitrarily many species to stably coexist in a sufficiently large system, despite strongly disordered competitive interactions. We analytically characterize the noise-induced transition to coexistence, showing that spatiotemporal noise drives power-law abundance fluctuations, leading to an anomalous scaling of moments known empirically as Taylor's law. At the metacommunity level, this manifests as an emergent sublinear self-inhibition that stabilizes diversity and renders the interaction disorder irrelevant in the high-diversity limit. Spatiotemporal noise thus provides a novel resolution to the diversity-stability paradox and a generic mechanism by which complex communities can persist.

q-bio.PE↗

arXiv2025

Learning to generalize in evolution through annealed population heterogeneity

Evolutionary systems must learn to generalize, often extrapolating from a limited set of selective conditions to anticipate future environmental changes. The mechanisms enabling such generalization remain poorly understood, despite their importance to predict ecological robustness, drug resistance, or design future-proof vaccination strategies. Here, we demonstrate that annealed population heterogeneity, wherein distinct individuals in the population experience different instances of a complex environment over time, can act as a form of implicit regularization and facilitate evolutionary generalization. Mathematically, annealed heterogeneity introduces a variance-weighted demographic noise term that penalizes across-environment fitness variance and effectively rescales the population size, thereby biasing evolution toward generalist solutions. This process is indeed analogous to a variant of the mini-batching strategy employed in stochastic gradient descent, where an effective multiplicative noise produces an inductive bias by triggering noise-induced transitions. Through numerical simulations and theoretical analysis we discuss the conditions under which variation in how individuals experience environmental selection can naturally promote evolutionary strategies that generalize across environments and anticipate novel challenges.

q-bio.PE↗

arXiv2025

Minimal model of self-organized clusters with phase transitions in ecological communities

In complex ecological communities, species may self-organize into clusters or clumps where highly similar species can coexist. The emergence of such species clusters can be captured by the interplay between neutral and niche theories. Based on the generalized Lotka-Volterra model of competition, we propose a minimal model for ecological communities in which the steady states contain self-organized clusters. In this model, species compete only with their neighbors in niche space through a common interaction strength. Unlike many previous theories, this model does not rely on random heterogeneity in interactions. Even in this minimal model where only the common interaction strength is varied, we find an exponentially large set of states that exhibit a rich variety of cluster patterns with different sizes and combinations. There are sharp phase transitions into the formation of clusters. There are also multiple phase transitions between different sets of possible cluster patterns, many of which accumulate near a small number of critical points. We analyze this phase structure using both numerical and analytical methods. In addition, the special case with only nearest neighbor interactions is exactly solvable using the method of transfer matrices from statistical mechanics. We analyze the critical behavior of these systems.

cond-mat.stat-mech↗

arXiv2025

Universal scaling of segment fluctuations in polymer and chromatin dynamics

We demonstrate how center-of-mass (COM) motion influences polymer segment fluctuations. Cancellation of internal forces, together with spatially uncorrelated external noise, generally yields COM diffusivity scaling as $1/s$ with segment length $s$, regardless of fractal dimension, viscoelasticity, or activity. This introduces distinct dynamic scaling corrections to two-point fluctuations and quenched-induced tangential correlations, validated by theory, simulations, and chromatin imaging data. In the latter, the extracted dynamic exponent reveals topological constraints, thereby resolving the discrepancy between chromatin's crumpled structure and its Rouse-like dynamics.

cond-mat.soft↗

arXiv2024

New sector morphologies emerge from anisotropic colony growth

Competition during range expansions is of great interest from both practical and theoretical view points. Experimentally, range expansions are often studied in homogeneous Petri dishes, which lack spatial anisotropy that might be present in realistic populations. Here, we analyze a model of anisotropic growth, based on coupled Kardar-Parisi-Zhang and Fisher-Kolmogorov-Petrovsky-Piskunov equations that describe surface growth and lateral competition. Compared to a previous study of isotropic growth, anisotropy relaxes a constraint between parameters of the model. We completely characterize spatial patterns and invasion velocities in this generalized model. In particular, we find that strong anisotropy results in a distinct morphology of spatial invasion with a kink in the displaced strain ahead of the boundary between the strains. This morphology of the out-competed strain is similar to a shock wave and serves as a signature of anisotropic growth.

nlin.PS↗

arXiv2024

A Minimal Framework for Optimizing Vaccination Protocols Targeting Highly Mutable Pathogens

A persistent public health challenge is finding immunization schemes that are effective in combating highly mutable pathogens such as HIV and influenza viruses. To address this, we analyze a simplified model of affinity maturation, the Darwinian evolutionary process B cells undergo during immunization. The vaccination protocol dictates selection forces that steer affinity maturation to generate antibodies. We focus on determining the optimal selection forces exerted by a generic time-dependent vaccination protocol to maximize production of broadly neutralizing antibodies (bnAbs) that can protect against a broad spectrum of pathogen strains. The model lends itself to a path integral representation and operator approximations within a mean-field limit, providing guiding principles for optimizing time-dependent vaccine-induced selection forces to enhance bnAb generation. We compare our analytical mean-field results with the outcomes of stochastic simulations and discuss their similarities and differences.

q-bio.PE↗

arXiv2024

Delayed excitations induce polymer looping and coherent motion

We consider inhomogeneous polymers driven by energy-consuming active processes which encode temporal patterns of athermal kicks. We find that such temporal excitation programs, propagated by tension along the polymer, can effectively couple distinct polymer loci. Consequently, distant loci exhibit correlated motions that fold the polymer into specific conformations, as set by the local actions of the active processes and their distribution along the polymer. Interestingly, active kicks that are canceled out by a time-delayed echo can induce strong compaction of the active polymer.

cond-mat.soft↗

arXiv2023

Universal characterization of epitope immunodominance from a multi-scale model of clonal competition in germinal centers

We introduce a novel, multi-scale model for affinity maturation, which aims to capture the intra-clonal, inter-clonal and epitope-specific organization of the B cell population in a germinal center. We describe the evolution of the B cell population via a quasispecies dynamics, with species corresponding to unique B cell receptors (BCRs), where the desired multi-scale structure is reflected on the mutational connectivity of the accessible BCR space, and on the statistical properties of its fitness landscape. Within this mathematical framework, we study the competition among classes of BCRs targeting different antigen epitopes, and construct an effective \emph{immunogenic space} where epitope immunodominance relations can be universally characterized. We finally study how varying the relative composition of a mixture of antigens with variable and conserved domains allows for a parametric exploration of this space, and identify general principles for the rational design of two-antigen cocktails.

q-bio.PE↗

arXiv2023

Inclusions, Boundaries and Disorder in Scalar Active Matter

Active systems are driven out of equilibrium by exchanging energy and momentum with their environment. This endows them with anomalous mechanical properties that we review in this colloquium for the case of dry scalar active matter, which has attracted considerable attention. These unusual properties lead to a rich physics when active fluids are in contact with boundaries, inclusions, tracers, or disordered potentials. Indeed, studies of the mechanical pressure of active fluids and of the dynamics of passive tracers have shown that active systems impact their environment in non-trivial ways, for example, by propelling and rotating anisotropic inclusions. Conversely, the long-ranged density and current modulations induced by localized obstacles show how the environment can have a far-reaching impact on active fluids. This is best exemplified by the propensity of bulk and boundary disorder to destroy bulk phase separation in active matter, showing active systems to be much more sensitive to their surroundings than passive ones. This colloquium aims at providing a unifying perspective on the rich interplay between active systems and their environments.

cond-mat.soft↗

arXiv2023

Scale-dependent heat transport in dissipative media via electromagnetic fluctuations

We develop a theory for heat transport via electromagnetic waves inside media, and use it to derive a spatially nonlocal thermal conductivity tensor, in terms of the electromagnetic Green's function and potential, for any given system. While typically negligible for optically dense bulk media, the electromagnetic component of conductivity can be significant for optically dilute media, and shows regimes of Fourier transport as well as unhindered transport. Moreover, the electromagnetic contribution is relevant even for dense media, when in presence of interfaces, as exemplified for the in-plane conductivity of a nanosheet, which shows a variety of phenomena, including absence of a Fourier regime.

physics.class-ph↗

arXiv2023

Escaping kinetic traps using non-reciprocal interactions

Kinetic traps are a notorious problem in equilibrium statistical mechanics, where temperature quenches ultimately fail to bring the system to low energy configurations. Using multifarious self-assembly as a model system, we introduce a mechanism to escape kinetic traps by utilizing non-reciprocal interactions between components. Introducing non-equilibrium effects offered by broken action-reaction symmetry in the system, we can push the trajectory of the system out of arrested dynamics. The dynamics of the model is studied using tools from the physics of interfaces and defects. Our proposal can find applications in self-assembly, glassy systems and systems with arrested dynamics.

cond-mat.stat-mech↗

arXiv2023

Competition on the edge of an expanding population

In growing populations, the fate of mutations depends on their competitive ability against the ancestor and their ability to colonize new territory. Here we present a theory that integrates both aspects of mutant fitness by coupling the classic description of one-dimensional competition (Fisher equation) to the minimal model of front shape (KPZ equation). We solved these equations and found three regimes, which are controlled solely by the expansion rates, solely by the competitive abilities, or by both. Collectively, our results provide a simple framework to study spatial competition.

q-bio.PE↗