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Sriram Ramaswamy

Publications and source records attributed to Sriram Ramaswamy.

At least 55 records · Page 3Linked to original sources

Origins and diagnostics of the nonequilibrium character of active systems

We present in detail a Langevin formalism for constructing stochastic dynamical equations for active-matter systems coupled to a thermal bath. We apply the formalism to clarify issues of principle regarding the sources and signatures of nonequilibrium behaviour in a variety of polar and apolar single-particle systems and polar flocks. We show that distance from thermal equilibrium depends on how time-reversal is implemented and hence on the reference equilibrium state. We predict characteristic forms for the frequency-resolved entropy production for an active polar particle in a harmonic potential, which should be testable in experiments.

cond-mat.soft↗

Kepler orbits of settling discs

The collective dynamics of objects moving through a viscous fluid is complex and counterintuitive. A key to understanding the role of nontrivial particle shape in this complexity is the interaction of a pair of sedimenting spheroids. We report experimental results on two discs settling at negligible Reynolds number ($\simeq 10^{-4}$), finding two classes of bound periodic orbits, each with transitions to scattering states. We account for these dynamics, at leading far-field order, through an effective Hamiltonian in which gravitational driving endows orientation with the properties of momentum. This leads to a precise correspondence with the Kepler problem of planetary motion for a wide range of initial conditions, and also to orbits with no Keplerian analogue. This notion of internal degrees of freedom manifesting themselves as an effective inertia is potentially a more general tool in Stokesian driven systems.

cond-mat.soft↗

Stability from activity

Suspensions of actively driven anisotropic objects exhibit distinctively nonequilibrium behaviors, and current theories predict that they are incapable of sustaining orientational order at high activity. By contrast, here we show that nematic suspensions on a substrate can display order at arbitrarily high activity due to a previously unreported, potentially stabilizing active force. The resulting nonequilibrium ordered phase displays robust giant number fluctuations that cannot be suppressed even by an incompressible solvent. Our results apply to virtually all experimental assays used to investigate the active nematic ordering of self-propelled colloids, bacterial suspensions and the cytoskeleton, and have testable implications in interpreting their nonequilibrium behaviors

cond-mat.soft↗

The low noise phase of a 2d active nematic

We consider a collection of self-driven apolar particles on a substrate that organize into an active nematic phase at sufficiently high density or low noise. Using the dynamical renormalization group, we systematically study the 2d fluctuating ordered phase in a coarse-grained hydrodynamic description involving both the nematic director and the conserved density field. In the presence of noise, we show that the system always displays only quasi-long ranged orientational order beyond a crossover scale. A careful analysis of the nonlinearities permitted by symmetry reveals that activity is dangerously irrelevant over the linearized description, allowing giant number fluctuations to persist though now with strong finite-size effects and a non-universal scaling exponent. Nonlinear effects from the active currents lead to power law correlations in the density field thereby preventing macroscopic phase separation in the thermodynamic limit.

cond-mat.soft↗

Active matter

The study of systems with sustained energy uptake and dissipation at the scale of the constituent particles is an area of central interest in nonequilibrium statistical physics. Identifying such systems as a distinct category -- Active Matter -- unifies our understanding of autonomous collective move- ment in the living world and in some surprising inanimate imitations. In this article I present the Active Matter framework, briefly recall some early work, review our recent results on single-particle and collective behaviour, including experiments on active granular monolayers, and discuss new directions for the future.

cond-mat.stat-mech↗

Hydrodynamic instabilities in active cholesteric liquid crystals

We describe the basic properties and consequences of introducing active stresses, with principal direction along the local director, in cholesteric liquid crystals. The helical ground state is found to be linearly unstable to extensile stresses, without threshold in the limit of infinite system size, whereas contractile stresses are hydrodynamically screened by the cholesteric elasticity to give a finite threshold. This is confirmed numerically and the non-linear consequences of instability, in both extensile and contractile cases, are studied. We also consider the stresses associated to defects in the cholesteric pitch ($λ$ lines) and show how the geometry near to the defect generates threshold-less flows reminiscent of those for defects in active nematics. At large extensile activity $λ$ lines are spontaneously created and can form steady state patterns sustained by constant active flows.

cond-mat.soft↗

Asymmetric exchange in flocks

As the constituents of a flock are polar, one expects a fore-aft asymmetry in their interactions. We show here that the resulting antisymmetric part of the "exchange coupling" between a bird and its neighbours, if large enough, destabilizes the flock through spontaneous turning of the birds. The same asymmetry also yields a natural mechanism for a difference between the speed of advection of information along the flock and the speed of the flock itself. We show that the absence of detailed balance, and not merely the breaking of Galilean invariance, is responsible for this difference. We delineate the conditions on parameters and wavenumber for the existence of the turning instability. Lastly we present an alternative perspective based on flow-alignment effects in an active liquid crystal with turning inertia in contact with a momentum sink.

cond-mat.stat-mech↗

Sorting motile rods by activity

We show, through experiments and simulations, that geometrically polar granular rods, rendered active by the transduction of vertical vibration, undergo a collective trapping phase transition in the presence of a V-shaped obstacle when the opening angle drops below a threshold value. We propose a mechanism that accounts qualitatively for the transition, based on the cooperative reduction of angular noise with increasing area fraction. We exploit the sensitivity of trapping to the persistence of directed motion to sort particles based on the statistical properties of their activity.

cond-mat.soft↗

Anisotropic Isometric Fluctuation Relations in experiment and theory on a self-propelled rod

The Isometric Fluctuation Relation (IFR) [P.I. Hurtado et al., PNAS 108, 7704 (2011)] relates the relative probability of current fluctuations of fixed magnitude in different spatial directions. We test its validity in an experiment on a tapered rod, rendered motile by vertical vibration and immersed in a sea of spherical beads. We analyse the statistics of the velocity vector of the rod and show that they depart significantly from the IFR of Hurtado et al. Aided by a Langevin-equation model we show that our measurements are largely described by an anisotropic generalization of the IFR [R. Villavicencio et al., EPL 105, 30009 (2014)], with no fitting parameters, but with a discrepancy in the prefactor whose origin may lie in the detailed statistics of the microscopic noise. The experimentally determined Large-Deviation Function of the velocity vector has a kink on a curve in the plane.

cond-mat.soft↗

Aspects of the density field in an active nematic

Active nematics are conceptually the simplest orientationally ordered phase of self-driven particles, but have proved to be a perennial source of surprises. We show here through numerical solution of coarse-grained equations for order parameter and density that the growth of the active nematic phase from the isotropic phase is necessarily accompanied by a clumping of the density. The growth kinetics of the density domains is shown to be faster than the 1/3-law expected for variables governed by a conservation law. Other results presented include the suppression of density fluctuations in the stationary ordered nematic by the imposition of an orienting field. We close by posing some open questions.

cond-mat.soft↗

Silent Flocks

Experiments find coherent information transfer through biological groups on length and time scales distinctly below those on which asymptotically correct hydrodynamic theories apply. We present here a new continuum theory of collective motion coupling the velocity and density fields of Toner and Tu to the inertial spin field recently introduced to describe information propagation in natural flocks of birds. The long-wavelength limit of the new equations reproduces Toner-Tu theory, while at shorter wavelengths (or, equivalently, smaller damping), spin fluctuations dominate over density fluctuations and second sound propagation of the kind observed in real flocks emerges. We study the dispersion relation of the new theory and find that when the speed of second sound is large, a gap sharply separates first from second sound modes. This gap implies the existence of `silent' flocks, namely medium-sized systems across which neither first nor second sound can propagate.

cond-mat.stat-mech↗

Flocking at a distance in active granular matter

The self-organised motion of vast numbers of creatures in a single direction is a spectacular example of emergent order. We recreate this phenomenon using actuated non-living components. We report here that millimetre-sized tapered rods, rendered motile by contact with an underlying vibrated surface and interacting through a medium of spherical beads, undergo a phase transition to a state of spontaneous alignment of velocities and orientations above a threshold bead area fraction. Guided by a detailed simulation model, we construct an analytical theory of this flocking transition, with two ingredients: a moving rod drags beads; neighbouring rods reorient in the resulting flow like a weathercock in the wind. Theory and experiment agree on the structure of our phase diagram in the plane of rod and bead concentrations and power-law spatial correlations near the phase boundary. Our discovery suggests possible new mechanisms for the collective transport of particulate or cellular matter.

cond-mat.soft↗

Activating membranes

We present a general dynamical theory of a membrane coupled to an actin cortex containing polymerizing filaments with active stresses and currents, and demonstrate that active membrane dynamics [Phys. Rev. Lett \textbf{84}, 3494 (2000)] and spontaneous shape oscillations emerge from this description. We also consider membrane instabilities and patterns induced by the presence of filaments with polar orientational correlations in the tangent plane of the membrane. The dynamical features we predict should be seen in a variety of cellular contexts involving the dynamics of the membrane-cytoskeleton composite and cytoskeletal extracts coupled to synthetic vesicles.

physics.bio-ph↗

Clusters, asters and collective oscillations in chemotactic colloids

The creation of synthetic systems that emulate the defining properties of living matter, such as motility, gradient-sensing, signalling and replication, is a grand challenge of biomimetics. Such imitations of life crucially contain active components that transform chemical energy into directed motion. These artificial realizations of motility point in the direction of a new paradigm in en- gineering, through the design of emergent behavior by manipulating properties at the scale of the individual components. Catalytic colloidal swimmers are a particularly promising example of such systems. Here we present a comprehensive theoretical description of gradient-sensing of an individual swimmer, leading controllably to chemotactic or anti-chemotactic behavior, and use it to construct a framework for studying their collective behavior. We find that both the positional and the orientational degrees of freedom of the active colloids can exhibit condensation, signalling formation of clusters and asters. The kinetics of catalysis introduces a natural control parameter for the range of the interaction mediated by the diffusing chemical species. For various regimes in parameter space in the long-ranged limit our system displays precise analogs to gravitational collapse, plasma oscillations and electrostatic screening. We present prescriptions for how to tune the surface properties of the colloids during fabrication to achieve each type of behavior.

cond-mat.soft↗

The Phase Synchronized State of Oriented Active Fluids

We present a theory for self-driven fluids, such as motorized cytoskeletal extracts or bacterial suspensions, that takes into account the underlying periodic duty cycle carried by the active particles of which the system is composed. We show that the fluid can enter a state in which the constituent elements synchronize their phases. This spontaneous breaking of time-translation invariance gives rise to flow instabilities distinct from those arising in phase-incoherent active matter.

cond-mat.soft↗

Actomyosin contractility rotates the cell nucleus

The nucleus of the eukaryotic cell functions amidst active cytoskeletal filaments, but its response to the stresses carried by these filaments is largely unexplored. We report here the results of studies of the translational and rotational dynamics of the nuclei of single fibroblast cells, with the effects of cell migration suppressed by plating onto fibronectin-coated micro-fabricated patterns. Patterns of the same area but different shapes and/or aspect ratio were used to study the effect of cell geometry on the dynamics. On circles, squares and equilateral triangles, the nucleus undergoes persistent rotational motion, while on high-aspect-ratio rectangles of the same area it moves only back and forth. The circle and the triangle showed respectively the largest and the smallest angular speed. We show that our observations can be understood through a hydrodynamic approach in which the nucleus is treated as a highly viscous inclusion residing in a less viscous fluid of orientable filaments endowed with active stresses. Lowering actin contractility selectively by introducing blebbistatin at low concentrations drastically reduced the speed and persistence time of the angular motion of the nucleus. Time-lapse imaging of actin revealed a correlated hydrodynamic flow around the nucleus, with profile and magnitude consistent with the results of our theoretical approach. Coherent intracellular flows and consequent nuclear rotation thus appear to be a generic property that cells must balance by specific mechanisms in order to maintain nuclear homeostasis.

physics.bio-ph↗

The glass susceptibility: growth kinetics and saturation under shear

We study the growth kinetics of glassy correlations in a structural glass by monitoring the evolution, within mode-coupling theory, of a suitably defined three-point function $χ_C(t,t_w)$ with time $t$ and waiting time $t_w$. From the complete wave vector-dependent equations of motion for domain growth we pass to a schematic limit to obtain a numerically tractable form. We find that the peak value $χ_C^P$ of $χ_C(t,t_w)$, which can be viewed as a correlation volume, grows as $t_w^{0.5}$, and the relaxation time as $t_w^{0.8}$, following a quench to a point deep in the glassy state. These results constitute a theoretical explanation of the simulation findings of Parisi [J. Phys. Chem. B {\bf 103}, 4128 (1999)] and Kob and Barrat [Phys. Rev. Lett. {\bf 78}, 4581 (1997)] and are also in qualitative agreement with Parsaeian and Castillo [Phys. Rev. E {\bf 78}, 060105(R) (2008)]. On the other hand, if the quench is to a point on the {\em liquid side}, the correlation volume grows to saturation. We present a similar calculation for the growth kinetics in a $p$-spin spin glass mean-field model where we find a slower growth, $χ_C^P \sim t_w^{0.13}$. Further, we show that a shear rate $\gdot$ cuts off the growth of glassy correlations when $t_w\sim 1/\gdot$ for quench in the glassy regime and $t_w=\min(t_r,1/\gdot)$ in the liquid, where $t_r$ is the relaxation time of the unsheared liquid. The relaxation time of the steady state fluid in this case is $\propto \gdot^{-0.8}$.

cond-mat.soft↗

Universal power law in crossover from integrability to quantum chaos

We study models of interacting fermions in one dimension to investigate the crossover from integrability to non-integrability, i.e., quantum chaos, as a function of system size. Using exact diagonalization of finite-sized systems, we study this crossover by obtaining the energy level statistics and Drude weight associated with transport. Our results reinforce the idea that for system size $L \to \infty$ non-integrability sets in for an arbitrarily small integrability-breaking perturbation. The crossover value of the perturbation scales as a power law $\sim L^{-3}$ when the integrable system is gapless and the scaling appears to be robust to microscopic details and the precise form of the perturbation. We conjecture that the exponent in the power law is characteristic of the random matrix ensemble describing the non-integrable system. For systems with a gap, the crossover scaling appears to be faster than a power law.

cond-mat.str-el↗